Nephridia are invertebrate organs which function similarly to kidneys. They remove metabolic wastes from an animal's body. They are present in many different invertebrate lines. There are two basic types, metanephridia and protonephridia.
Metanephridia
A
metanephridium (pl.
metanephridia) is a type of
excretory gland or
nephridium found in many types of
invertebrates such as
annelids,
arthropods and
mollusks. It typically consists of a ciliated funnel opening into the body cavity or
coelom connected to a
duct which may be variously glandularized, folded or expanded (vesiculate) and which typically opens to the
organism's exterior. These ciliated tubules pump water carrying surplus
ions,
metabolic waste,
toxins from
food, and useless
hormones out of the organism through openings known as
nephrostomes. The primary
urine produced by
filtration of blood (or a similar functioning component) are modified into secondary urine through selective
reabsorption by the
cells lining the metanephridium.
In many earthworms the nephridial ducts open into the digestive tract instead, a condition known as enteronephry.
Protonephridia
A
protonephridium is a network of dead-end
tubules lacking internal openings. The ends are called "
flame bulbs", they function in
osmoregulation. The terminal cells are located in the bottom of the protonephridium. Each cell has one or several
cilia and when they beat inside the protonephridial tube they create an outward going current and hence a partial
vacuum in the bottom of the tube. Because of this partial vacuum
waste fluids from the inside out the animal are pulled through small perforations in the terminal cells and into the protonephridium. The perforations in the terminal cell are so large that small
molecules can pass whereas larger
proteins are retained in the animal. From the bottom of the protonephridium the waste fluid is led through the tube, formed by the canal cells, and exits the animal from a small opening formed by the
nephridiopore. Protonephridia are found in simpler organisms such as
flatworms.
原肾管
Animal anatomy | Flatworms
Annelids | Nephridium