The bottom quark is a third-generation quark with a charge of -(1/3)e. Although all quarks are described in a similar way by the theory of quantum chromodynamics, the bottom quark's large mass (around 4 GeV, or four times the mass of a proton) gives it a distinctive signature that makes it relatively easy to identify experimentally (using a technique called B-tagging). Because three generations of quark are required for CP violation (see CKM matrix for an explanation of why), mesons containing the bottom quark are the easiest particles to use to investigate the phenomenon; this is being done at the BaBar and Belle experiments. The bottom quark is also notable because it is a product in almost all top quark decays, and a frequent decay product for the hypothetical Higgs boson if it is sufficiently light.
The bottom quark was discovered at Fermilab in 1977. On its discovery, a valiant effort was made by many particle physicists to name it "Beauty", paired along with "Truth". Poetic license gave way to pragmatics, and the quarks were named "top" and "bottom" instead.
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