Cochlospermaceae is a family of 20-25 species distributed among two genera of trees and shrubs. They occur widely throughout the tropical regions of the world, but are curiously absent from Malaysia. Most species in this family are mesophytic or xerophytic, growing primarily in drier climates.
Leaf form and anatomy
Leaves
alternate;
petiolate; non-sheathing; gland-dotted, or not gland-dotted;
aromatic, or without marked odor; simple.
Lamina dissected;
palmatifid;
palmately veined; cross-venulate. Leaves
stipulate.
Lamina margins
serrate. Leaves without a persistent basal
meristem. Mucilaginous
epidermis often present, minor leaf veins without
phloem transfer cells (
Cochlospermum).
Stem anatomy
Secretory cavities present (in the
cortex and
pith); with
mucilage. Cortical bundles absent.
Medullary bundles absent. Internal
phloem absent. Secondary thickening developing from a conventional
cambial ring. The secondary
phloem stratified into hard (fibrous) and soft (parenchymatous) zones.
Xylem with fibre
tracheids.
Vessels without vestured pits (but with vestured fibre pits).
Wood partially storied (VPI), or not storied;
parenchyma predominantly apotracheal.
Inflorescence, floral, fruit and seed morphology
Flowers aggregated in
inflorescences; in
panicles and in
racemes. Flowers large; regular to somewhat irregular. Free hypanthium absent.
Perianth
Perianth with distinct
calyx and
corolla.
Calyx 4 or 5-lobed; 1 whorled;
polysepalous (multiple distinct sepals); not persistent (deciduous);
imbricate.
Corolla 4 or 5-lobed; 1 whorled;
polypetalous (multiple distinct petals);
imbricate, or
contorted; yellow.
Androecium
Androecium of 15-60 exclusively fertile
stamens, maturing centrifugally; free of the
perianth. Stamens equal to markedly unequal, free of one another.
Anthers dehiscing via pores (these
apical, often
confluent).
Pollen grains 3-
aperturate;
colporate; 2-celled.
Gynoecium
Gynoecium 3–5 carpelled. The
pistil 1 or 3 celled.
Gynoecium syncarpous, superior. Ovary 1 or 3 locular. Styles 1;
attenuate from the ovary;
apical. Stigmas 1 (minute).
Placentation when
unilocular,
parietal; when
trilocular,
axile. Ovules in the single cavity when unilocular, 20–100 or more; when trilocular, 20–50 per locule;
anatropous;
bitegmic;
crassinucellate. Outer
integument contributing to the
micropyle.
Endosperm formation nuclear.
Fruits and seeds
Fruit non-fleshy;
dehiscent; a
capsule. Capsules three to five valvular.
Seeds endospermic.
Endosperm oily. Embryo straight, or curved, or coiled.
Micropyle zigzag.
Physiology and biochemistry
Proanthocyanidins present;
cyanidin.
Flavonols present;
kaempferol,
quercetin, and
myricetin.
Ellagic acid present (
Cochlospermum).
Saponins/
sapogenins present.
Aluminium accumulation not found.
External links
Cochlospermaceae | Cochlospermaceae | Cochlospermaceae