Amphicoelias (correctly pronounced AM-fi-KOI-lee-as, commonly pronounced AM-fi-SEEL-ee-as, meaning "double cavities") was a genus of sauropod dinosaur, which may have included the largest dinosaur ever discovered (A. fragillimus). If it truly existed, A. fragillimus would be the longest vertebrate by a considerable margin, and it would have a mass that rivals the heaviest animal known, the blue whale. However, because the only fossil evidence for its existence was lost shortly after been found in the 1870s, evidence survives only in drawings, and doubt has been expressed about their veracity.
The neural arch was packaged and sent by train to a Museum in New York, but unfortunately it never arrived; apparently it crumbled to dust while en route. The only evidence remaining of its existence is the description and a line drawing in a scientific journal.
Marsh was winning the bone war. He discovered a total of 86 new species during their decades-long rivalry, due in part to his discovery of the Como Bluff site, near Medicine Bow, Wyoming, one of the richest sources of fossils ever uncovered. Cope only discovered 56; many of the fossils he unearthed were of species that had already been named, or were of uncertain origin, so he was not credited with their discovery. As a result, Marsh's name is attached to many of the most famous dinosaurs, such as Triceratops, Allosaurus, Diplodocus, and Stegosaurus, while even Cope's most famous discoveries, such as Dimetrodon (not actually a dinosaur), Camarasaurus, Coelophysis, and Monoclonius are more obscure. So while there is no evidence of falsification, the circumstances surrounding Amphicoelias do raise some doubt about the authenticity of the find.
By comparison the blue whale, which is the longest living creature, only reaches 30–33 m (100–110 ft.) in length, and the longest dinosaur for which solid evidence exists, Supersaurus, is estimated to be only about 40 m (130 ft.) long.
While A. fragillimus was relatively thin, its enormous size still makes it very massive. The heaviest blue whale on record weighed about 175 t (195 tons), which is just slightly larger than the upper range of estimates for Amphicoelias. For the sake of comparison, the heaviest dinosaur that we have good evidence for, the Argentinosaurus, only weighed 80–100 t (90–100 tons), although if the size estimates can be validated, it would still be lighter than Bruhathkayosaurus, which is estimated to weight 175–220 t (190–240 tons) (based on a poor description of a partial fossil).
The genus Amphicoelias is poorly known. The first named species in the genus, Amphicoelias altus (holotype specimen AMHD 5764), was discovered by Cope the year before, in 1877. But while it is only represented by a partial skeleton, there are enough diagnostic characteristics to provisionally define the genus. A. altus is known from slightly better remains, but is smaller than A. fragillimus. McIntosh (1998, Modern Geology 23:481-506) stated in 1998 that there is no reason to suspect the two are different species. "A. fragillimus" is probably just a very large individual of A. altus.
Diplodocoids | Jurassic dinosaurs | North American dinosaurs
Amphicoelias | アンフィコエリアス | Amphicoelias fragillimus | Amphicoelias
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